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According to Wikipedia (search Zucchetto) the origins seem to be largely practical - to keep the priests head warm in cold churches. Tradition still survives for bishops to remove the Zucchetto when coming into contact with the Blessed Sacrament (and churchgoers will see the Bishop remove the Zucchetto at the beginning of the Eucharistic prayers {more specifically, the Preface} until just before the Final Prayer) - Bishops also remove the Zucchetto for benediction, or for processions with the Blessed Sacrament, such as on Holy Thursday or on Corpus Christi).

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Catholic AnswerRegardless of its orgins, the Zuchetto (the small skull cap) is part of the attire specified for Bishops, Archbishops, Cardinals, and the Pope. It is a sign of their Office and the color is different for each. An abbot, is required to wear a black Zuchetto, and priests may wear a black one, as well, but it is not specified.
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Q: Why does a Catholic bishop wear a Zuchetto?
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