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It depends on use.

The Wizard decreed: There will be NO sharing of shoe laces. All those who participate in this dastardly act will be forced to squish noodles through their teeth. Here, "all those" is plural, so the plural form of the verb is used.

The investigator found irrefutable evidence that a large group of people gathered there and shared bellybutton lint. "Who participates in such heinous things?" The investigator wondered, mystified. Here the use of the word "who" is singular, as in "what sort of individual participates". So the singular form of the verb is used.

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12y ago
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6mo ago

The word "who" is used as a singular pronoun, so it should be followed by the singular form "participates." For example, "Who participates in the event?"

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Q: Who participate or who participates?
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