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The event which historians consider as marking the end of the western part of the Roman Empire and the beginning of the Middle Ages (not dark ages) is the deposition of Romulus Augustus, the last emperor of the western part of the empire in 476.
Yes, polyphony was used during the Middle Ages. We do not have any real idea of when polyphony was introduced, but the earliest texts describing it date to the Early Middle Ages. By the end of the Middle Ages, it was highly developed. There is a link below to an article on polyphony, beginning at the section on its origins.
The next age after the Middle Ages was the Renaissance or Early Modern Age. There is a bit of overlap between the two, as the Renaissance is taken as beginning about 1350 and the Middle Ages ending about 1450 or so.
The Middle Ages began in the 5th century and end in the 15th century.