A closeup of the front of a human skull.
The skull is a bony structure found in many animals which serves as the general framework for the head. The
skull supports the structures of the face and protects the head
against injury.
The skull can be subdivided into two parts: the cranium and the mandible. A skull that is missing a mandible is
only a cranium; this is the source of a very commonly made error in terminology. Those animals having skulls are called
craniates.
Protection of the brain is only one part of the function of a bony skull. For example, a fixed distance between the eyes is
essential for stereoscopic vision, and a fixed position for the ears helps the brain to use
auditory cues to judge direction and distance of sounds. In some animals, the skull also has a defensive function (e.g. horned
ungulates); the frontal bone is where horns are
mounted.
Human skulls
-
In humans, the adult skull is normally made up of 22 bones. Except for the
mandible, all of the bones of the skull are joined together by sutures, rigid articulations permitting very little movement. Eight bones form the neurocranium
(braincase), a protective vault surrounding the brain. Seventeen bones form the
skunt, the bones supporting the face. Encased within the temporal bones are the six
ear ossicles of the middle ears, though these are
not part of the skull. The hyoid bone, supporting the tongue,
is usually not considered as part of the skull either, as it does not articulate with any other bones, though it may be
considered a part of the skunt.
The skull contains the sinus cavities, which are air-filled cavities lined with
respiratory epithelium, which also lines the large airways. The exact functions
of the sinuses are unclear; they may contribute to lessening the weight of the skull with a minimal reduction in strength,or they
may be important in improving the resonance of the voice. In some animals, such as the elephant, the sinuses are extensive. The elephant skull needs to be very large, to form an attachment for
muscles of the neck and trunk, but is also unexpectedly light; the comparatively small brain-case is surrounded by large sinuses
which reduce the weight. The meninges are the three layers, or membranes, which surround the
structures of the nervous system. They are known as the dura
mater, the arachnoid mater and the pia mater.
Other than being classified together, they have little in common with each other.
In humans, the anatomical position for the skull is the Frankfurt plane, where the lower margins of the orbits and the
upper borders of the ear canals are all in a horizontal plane. This is the position where the
subject is standing and looking directly forward. For comparison, the skulls of other species, notably primates and hominids, may sometimes be studied in the Frankfurt plane.
However, this does not always equate to a natural posture in life.
Possible types of skull fractures
The protection of the brain and other vascular and respitory structures provided by the skull, can be compromised if the skull
is fractured. Various types of fractures are described below.
Mid-facial Skeletal fracture
The mid facial skeleton is made up of a considerable number of bones which are rarely, if ever, fractured in isolation
The structure is such that it is able to withstand considerable force from below, but the bones are easily fractured by
relatively trivial forces applied from other directions
Analogous to a ‘matchbox’ sitting below and in front of a hard shell containing the brain and differs quite markedly from the
rigid projection of the mandible below
Le Fort I Fractures
Low-level / Guerin type fractures
Horizontal fracture of the maxilla immediately above the teeth and palate
Piriform fossa across maxilla to pterygoid fissure
May occur as a single entity or in association with le fort II and III fractures
Sometimes present in association with a downwardly displaced fracture of the zygomatic
complex
Le Fort II Fractures
Pyramidal or suprazygomatic fractures
Fracture extends from dorsum of nose, across medial walls of orbit across the maxilla below
the zygomatic bone to the pterygomaxillary fissure
Le Fort III Fractures
High level or suprazygomatic fractures
The facial bones, including the zygomas are detached from the anterior cranial base
Fracture line extends from the dorsum of the nose and cribiform plate along the medial and up
the lateral wall of the orbit to the ZF suture
Animal skulls
Temporal Fenestra
The temporal fenestra are anatomical features of the amniote skull, characterised by bilaterally symmetrical holes (fenestrae) in the temporal bone. Depending on the
lineage of a given animal, two, one, or no pairs of temporal fenestrae may be present, above or below the postorbital and squamosal bones. The upper temporal fenestrae are also
known as the supratemporal fenestrae, and the lower temporal fenestrae are also known as the infratemporal fenestrae. The
presence and morphology of the temporal fenestra is critical for taxonomic classification of the synapsids, of which mammals are
part.
Physiological speculation associates it with a rise in metabolic rates and an increase in jaw musculature. The earlier
amniotes of the Carboniferous did not have temporal fenestrae but the more advanced sauropsids and synapsids did. As time
progressed, sauropsids' and synapsids' temporal fenestrae became more modified and larger to make stronger bites and more jaw
muscles. Dinosaurs, which are sauropsids, have large advanced openings and their descendants, the birds, have temporal fenestrae
which have been modified. Mammals, which are synapsids, possess no fenestral openings in the skull, as the trait has been
modified. They do, though, still have the temporal orbit (which resembles an opening) and the temporal muscles. It is a hole in
the head and is situated to the rear of the orbit behind the eye.
Classification
There are four types of amniote skull, classified by the number and location of their fenestra. These are:
- Anapsida - no openings
- Synapsida - one low opening (beneath the postorbital and squamosal bones)
- Euryapsida - one high opening (above the postorbital and squamosal bones); euryapsids
actually evolved from a diapsid configuration, losing their lower temporal fenestra.
- Diapsida - two openings
Evolutionary, they are related like so:
|
|
|
|
A coypu skull, a typical rodent
|
|
|
|
|
See also
References
- White, T.D. 1991. Human osteology. Academic Press, Inc. San Diego, CA.
External links
Wikimedia Commons has media related to:
This entry is from Wikipedia, the leading user-contributed encyclopedia. It may not have been reviewed by professional editors (see full disclaimer)